We are celebrating the seventy-fifth anniversary of Professor Pavel Štys’ birth

Miroslav Papáček, Carl W. Schaefer, Petr Kment

Four chapters about the monophyly of insect ‘orders’: A review of recent phylogenetic contributions

Jan Zrzavý

Recent phylogenetic analyses, both morphological and molecular, strongly support the monophyly of most insect ‘orders’. On the contrary, the Blattaria, Psocoptera, and Mecoptera are definitely paraphyletic (with respect of the Isoptera, Phthiraptera, and Siphonaptera, respectively), and the…

A new species of Systelloderes (Hemiptera: Heteroptera: Enicocephalidae) from South Africa

Petr Baňař

Systelloderes stysi sp. nov. from South Africa is described, and the apicitibial and tarsal armature of the fore leg is illustrated and described in detail.

Two new species of Kokeshia from India and Thailand (Hemiptera: Heteroptera: Schizopteridae)

Dávid Rédei

Kokeshia stysi sp. nov. (India) and K. oroszi sp. nov. (Thailand) (Hete-roptera: Schizopteridae) are described. A key for the identification of the known species of Kokeshia is presented.

Evolution of the semi-aquatic bugs (Hemiptera: Heteroptera: Gerromorpha) with a re-interpretation of the fossil record

Jakob Damgaard

The fossil history of semi-aquatic bugs (Gerromorpha) is reviewed in light of the many important recent records and new developments in our under-standing of the phylogeny of the group. Based on the age of its sister group, the Panheteroptera, the Gerromorpha probably extends back into the Triassic,…

Notes on aquatic and semiaquatic bugs (Hemiptera: Heteroptera: Nepomorpha, Gerromorpha) from Malesia with description of three new species

Ping-ping Chen, Nico Nieser, Ivor Lansbury

Micronecta lumutensis sp. nov. (Micronectidae) from Kalimantan (Borneo, Indonesia), Enithares rinjani sp. nov. (Notonectidae) from Lombok (Indonesia) and Neusterensifer stysi sp. nov. (Veliidae) from New Guinea are newly described. In addition, distributional records are given for three species of…

A new Indian Ocean species of Ochterus from the island of Mauritius (Hemiptera: Heteroptera: Ochteridae)

Dan A. Polhemus, John T. Polhemus

Ochterus stysi sp. nov. is described from the island of Mauritius in the western Indian Ocean. This species is only the second member of the family Ochteridae known from the islands of the Indian Ocean east of Madagascar, and the first recorded from the Mascarene Archipelago. Comparative notes are…

Intraspecific morphological polymorphism in Naucoridae (Hemiptera: Heteroptera) with notes on nomenclature and synonymy

John T. Polhemus, Dan A. Polhemus

Trends in intraspecific variation among certain aspects of pronotal and tarsal morphology within the family Naucoridae are reviewed, and linked to previously flawed taxonomic decisions regarding species separation or generic assignments within this group, particularly in the subfamily Laccocorinae.…

Structure and development of the reproductive system in Aphelocheirus aestivalis (Hemiptera: Heteroptera: Nepomorpha: Aphelocheiridae)

Miroslav Papáček, Tomáš Soldán

We study the histology, ultrastructure and morphological arrangement of male and female reproductive system of Aphelocheirus aestivalis (Fabricius, 1803) in the nymphs of instar 1-5 and variously aged apterous adults. The anlage of gonads becomes well apparent already in instar 1, testicular…

Notes on the endemic Philippine genus Orthosaldula (Hemiptera: Heteroptera: Saldidae), with descriptions of two new species

Herbert Zettel, Victor P. Gapud

The endemic Philippine genus Orthosaldula Gapud, 1986 and the type species O. rubroalata Gapud, 1986 from central and southern Luzon are redescribed. A male from Quezon Province, Central Luzon, is designated as the lectotype of O. rubroalata; the year of original publication is corrected. Two…

From four- to three-segmented labium in Reduviidae (Hemiptera: Heteroptera)

Christiane Weirauch

The majority of Reduviidae – unlike most other Heteroptera – has a labium that consists of only three segments. The first segment is said to be either lost or fused to the head capsule. Cladistic analysis shows that this loss or fusion occurred once or twice among basal Reduviidae and that the…

Two new species of the emesine assassin bug genus Ploiaria (Hemiptera: Heteroptera: Reduviidae) from Indonesia

Tadashi Ishikawa, Wayan Susila, Shûji Okajima

Two new species of the emesine reduviid genus Ploiaria Scopoli, 1786, P. stysi sp. nov. and P. paveli sp. nov., are described from Bali and Flores, Indonesia. Ploiaria stysi sp. nov. differs from P. maai Wygodzinsky, 1966, by the male antennal segments I and II lacking long setae, the procoxa 1.3…

Oncocephalus stysi, a new species of Stenopodainae (Hemiptera: Heteroptera: Reduviidae) from Israel

Pierre Moulet

A new species of the genus Oncocephalus Klug, 1830, O. stysi sp. nov., is described from Israel. The new species is very similar and related to O. aspericollis Reuter, 1882, and O. hierosolymensis Moulet, 2001. A new species-group, the aspericollis-group, is defined within Oncocephalus for these…

New records of Scotomedes alienus sikkimensis (Hemiptera: Heteroptera: Velocipedidae) from Nepal

Denise Wyniger

New records are given from Nepal for the scarce Scotomedes alienus sikkimensis van Doesburg, 2004 (Velocipedidae).

Fulvius stysi, a new species of Cylapinae (Hemiptera: Heteroptera: Miridae) from Papua New Guinea

Frédéric Chérot, Jacek Gorczyca

A new species of the genus Fulvius Stål, 1862, F. stysi sp. nov., is described from Baiteta Forest, Madang Province, Papua New Guinea. The male and female genital structures are illustrated.

Stysiofulvius, a new genus of Cylapinae (Hemiptera: Heteroptera: Miridae) from the Peninsular Malaysia

Jacek Gorczyca, Frédéric Chérot

Stysiofulvius gen. nov., a new genus, and S. hulinkai sp. nov., a new species of the plant bug subfamily Cylapinae are described from Malaysia. Illustrations of the dorsal habitus of the male and genitalic structures of both sexes are given.

A new species of the genus Phytocoris (Hemiptera: Heteroptera: Miridae)

Rauno E. Linnavuori

A new species of the family Miridae, Phytocoris (Compsocerocoris) stysi sp. nov., is described from the Khuzestan province in southern Iran.

Acetropis stysi, a new species from Spain (Hemiptera: Heteroptera: Miridae)

Reinhard Remane, Hannes Günther

Acetropis stysi sp. nov. is described from Sierra de Gredos of Ávila province in Central Spain. The new species is most similar to A. longirostris Puton, 1875, but in some respects is also similar to A. carinata (Herrich-Schaeffer, 1841). Males are macropterous, females macropterous and…

A review of the Far East Asian mirine plant bug genus Loristes (Hemiptera: Heteroptera: Miridae: Mirinae), with description of a new species from Japan

Tomohide Yasunaga

The little known mirine plant bug genus Loristes Josifov & Kerzhner, 1972 is reviewed. A species from Japan which was previously regarded conspecific with the continental L. decoratus (Reuter, 1908) is here described as L. paveli sp. nov.

Three new species of Phylini (Hemiptera: Heteroptera: Miridae: Phylinae) from Central Asia

Fedor V. Konstantinov

Three new species of Phylini, Pleuroxonotus stysi sp. nov. (Eastern Kazakhstan), Macrotylus subattenuatus sp. nov. (Northern Kazakhstan), and Lepidargyrus fasciatus sp. nov. (Uzbekistan) are described. Illustrations of the male genitalia, tarsus and pretarsus, photographs of the dorsal habitus,…

A new species of Megalocoleus (Hemiptera: Heteroptera: Miridae: Phylinae) from Morocco

Armand Matocq

Megalocoleus stysi sp. nov. is described from Morocco and its habitus and male and female genitalia are illustrated. An amendment to the key for Megalocoleus Reuter, 1890, is proposed.

Two new Phylinae (Hemiptera: Heteroptera: Miridae) from the Canary Islands

Jordi Ribes, Santiago Pagola-Carte, Ernst Heiss

Two new species of Phylinae (Hemiptera: Heteroptera: Miridae) are described from the Canary Islands. Their habitus, several characters of external morphology and genitalia are illustrated. Systellonotus stysi sp. nov., belonging to the tribe Hallodapini, is well recognized by its scutellar hump and…

Systematics, biogeography and host associations of the lace bug genus Inoma (Hemiptera: Heteroptera: Tingidae)

Gerasimos Cassis, Celia Symonds

The lace bug genus Inoma Hacker, 1927 is revised, including a redescription of the genus and the type species, I. multispinosa Hacker, 1927. Eight species are described as new to science, as follows: I. arrernte sp. nov., I. breviseta sp. nov., I. fuscata sp. nov., I. innamincka sp. nov., I.…

Phymacysta stysi, a new species of lace bug from Dominican amber (Hemiptera: Heteroptera: Tingidae: Tinginae)

Viktor B. Golub, Yuri A. Popov, Eric Guilbert

One new species of lace bug (Heteroptera, Tingidae), Phymacysta stysi sp. nov., is described from the Miocene Dominican amber, and its relationships are discussed. It belongs to the extant Neotropical genus Phymacysta Monte, 1942. A key of Phymacysta species including the new species and the fossil…

New data on the Tingidae from Gabon (Hemiptera: Heteroptera)

Eric Guilbert

Two species of the Tingidae, Belenus thomasi Drake, 1957, and Phatnoma maynei Schouteden, 1916, are recorded from Gabon for the first time, and one new species, Cysteochila stysi sp. nov., is described. The number of species of the Tingidae recently known from Gabon rises to 10.

Pavlostysia wunderlichi gen. nov. and sp. nov., the first fossil spider-web bug (Hemiptera: Heteroptera: Cimicomorpha: Plokiophilidae) from the Baltic Eocene amber

Yuri A. Popov

Pavlostysia wunderlichi gen. nov. and sp. nov., a remarkable new fossil genus and species of the cimicomorphan family Plokiophilidae, is described from Baltic amber.

Rajburicoris, a new genus of Cardiastethini, and discussion of the systematic position of Dufouriellus (Hemiptera: Heteroptera: Anthocoridae)

Diego L. Carpintero, Pablo M. Dellapé

Rajburicoris stysi gen. nov. and sp. nov. from Thailand is described and the male and female genitalia are examined and illustrated. A review of the diagnostic characters of the tribe Cardiastethini Carayon, 1972, nom. restit., is given. The systematic position of the controversial genus…

A pair of basi-abdominal sex pheromone glands in the male of some burrower bugs (Hemiptera: Heteroptera: Cydnidae)

Dominique Pluot-Sigwalt

Paired pleural basi-abdominal glands (PBA glands) were found in males of some burrower bugs (Pentatomoidea: Cydnidae). So far, they seem to be restricted to the subfamily Cydninae, tribes Cydnini (except Cydnus aterrimus (Forster, 1771) and Geotomini, and are absent in examined representatives of…

New subtribes and a new genus of Podopini (Heteroptera: Pentatomidae: Podopinae)

Dmitry A. Gapon

The tribe Podopini is divided into three subtribes: Kayesiina subtrib. nov., Podopina, and Scotinopharina subtrib. nov., differing in the length and shape of jugae, presence or absence of a tooth on the anterior part of bucculae and transverse carinae on pronotum, structure of pygophore, parandria,…

Two new species of the genus Chloropepla (Hemiptera: Pentatomidae: Pentatominae) from Brazil

Jocélia Grazia, Cristiano F. Schwertner, Caroline Greve

Chloropepla paveli sp. nov. and C. stysi sp. nov., from Brazil are described, based on the morphology of genitalia for both sexes. Prior to this study, the genus was represented in Amazon region as well as in southern and southeastern Brazil. The addition of the two species described in this paper…

A revision of the endemic Madagascan genus Triplatyx (Hemiptera: Heteroptera: Pentatomidae)

Petr Kment

The endemic Madagascan genus Triplatyx Horváth, 1904 (Pentatomoidea: Pentatomidae: Pentatominae: Triplatygini) is redescribed. Five species are recognized, two of them being new: T. bilobatus Cachan, 1952, T. dubius Jensen-Haarup, 1931, T. kerzhneri sp. nov., T. quadraticeps Horváth, 1904, and T.…

Massocephalus stysi, a new species of Pentatomidae (Hemiptera: Heteroptera) from the Philippines

David A. Rider

The pentatomid genus Massocephalus Dallas, 1851 and its only previously known species, M. maculatus Dallas, 1851 are redescribed. The tribal placement of Massocephalus is briefly discussed. One new species, M. stysi sp. nov. is described. A key to the species is provided.

A revision of the Entisberus group (Hemiptera: Heteroptera: Rhyparochromidae)

Elöd KONDOROSY

The genus Entisberus Distant, 1903, and related genera are revised. Neoentisberus Scudder, 1968, syn. nov., is synonymized with Retoka China, 1935, and Entisberus gibbus Zheng, 1981 is transferred to Retoka, which results in the following new combinations: Retoka brincki (Slater, 1979) comb. nov.,…

New taxa of the Largidae and Pyrrhocoridae (Hemiptera: Heteroptera) from the Oriental Region

Jaroslav L. Stehlík, Zdeněk Jindra

The following new taxa are described in the Largidae and Pyrrhocoridae: Largidae – Delacampius alboarcuatus sp. nov. (Indonesia: Bali), D. parvulus sp. nov. (Thailand), D. siberutensis sp. nov. (Indonesia: Siberut), Iphita fasciata sp. nov. (India: Maharastra), I. fuscorubra sp. nov. (India:…

Two new species of Anasa (Hemiptera: Heteroptera: Coreidae: Coreini) from Mexico and Central America

Harry Brailovsky, Ernesto Barrera

Two new species of Anasa Amyot & Serville, 1843 (Coreidae), Anasa paveli sp. nov. from Mexico, and A. stysi sp. nov. from Guatemala, Mexico, and Nicaragua, are described. Drawings of the pronotum, hemelytra, male genital capsule, and paramere are provided.

A review of the genus Satonius (Coleoptera: Myxophaga: Torridincolidae): taxonomic revision, larval morphology, notes on wing polymorphism, and phylogenetic implications

Jiří Hájek, Martin Fikáček

The genus Satonius Endrödy-Younga, 1997, is revised. Three new species are described from China: Satonius schoenmanni sp. nov. (Hunan province), S. stysi sp. nov. (Yunnan province), and S. wangi sp. nov. (Zhejiang province). Two additional new species from the Chinese provinces Fujian and Anhui are…

Taxonomic changes within the tribe Hyperini (Coleoptera: Curculionidae)

Jiří Skuhrovec

The taxonomic status of three genera (Donus Jekel, 1865, Brachypera Capiomont, 1868 stat. restit., and Hypera Germar, 1817) in the tribe Hyperini is clarified, including the identity of the type species of each genus, and problematic groups in each genus are commented. Neoglanis Alonso-Zarazaga &…

Tachysphex stysi sp. nov. (Hymenoptera: Apoidea: Crabronidae) from Central Asia

Jakub Straka

Tachysphex austriacus species subgroup of T. pompiliformis species group is defined. Tachysphex stysi sp. nov., a new member of this subgroup, is described. The new species is known from Kazakhstan, Tajikistan, and Turkmenistan.